State Key Laboratory of Stem Cell and Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences,National Laboratory of Biomacromolecules, Institute of Biophysics, Chinese Academy of Sciences,State Key Laboratory of Stem Cell and Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences,National Laboratory of Biomacromolecules, Institute of Biophysics, Chinese Academy of Sciences,National Laboratory of Biomacromolecules, Institute of Biophysics, Chinese Academy of Sciences

This work was supported by grants from The Strategic Priority Research Program of the Chinese Academy of Sciences (XDA16010100), The National Basic Research Program of China (2015CB964800, 2017YFA0103304, 2017YFA0102802, 2014CB910503, 2014CB964600, 2018YFA0107203), and the National Natural Science Foundation of China (91749202, 31471394, 31671429, 91749123, 81625009, 81330008, 81371342, 81471414, 81422017, 81601233, 81671377, 31601109, 31601158, 81771515, 81701388, 31571533, 31621004, 81822018)