College of Bioengineering,Tianjin University of Science Technology,Tianjin Key Laboratory of Industrial Microbiology,Ministry of Education Key Laboratory of Industrial Microbiology,College of Bioengineering,Tianjin University of Science Technology,Tianjin Key Laboratory of Industrial Microbiology,Ministry of Education Key Laboratory of Industrial Microbiology,College of Bioengineering,Tianjin University of Science Technology,Tianjin Key Laboratory of Industrial Microbiology,Ministry of Education Key Laboratory of Industrial Microbiology,College of Bioengineering,Tianjin University of Science Technology,Tianjin Key Laboratory of Industrial Microbiology,Ministry of Education Key Laboratory of Industrial Microbiology,College of Bioengineering,Tianjin University of Science Technology,Tianjin Key Laboratory of Industrial Microbiology,Ministry of Education Key Laboratory of Industrial Microbiology,College of Bioengineering,Tianjin University of Science Technology,Tianjin Key Laboratory of Industrial Microbiology,Ministry of Education Key Laboratory of Industrial Microbiology,College of Bioengineering,Tianjin University of Science Technology,Tianjin Key Laboratory of Industrial Microbiology,Ministry of Education Key Laboratory of Industrial Microbiology,College of Bioengineering,Tianjin University of Science Technology,Tianjin Key Laboratory of Industrial Microbiology,Ministry of Education Key Laboratory of Industrial Microbiology

This work was supported by grants from the Ministry of Science and Technology of China (2013DFG52490), Tianjin Municipal Science and Technology Commission (12ZXCXSY07500), The National Natural Science Fundation of China (21076157)