1.1)State Key Laboratory of Brain and Cognitive Science, Institute of Biophysics, Chinese Academy of Sciences, Beijing 100101, China;2.2)University of Chinese Academy of Sciences, Beijing 100049, China;3.3)Ruijin Hospital, Luwan Branch, Shanghai Jiao Tong University School of Medicine, Shanghai 200020, China;4.4)Institute of Automation, Chinese Academy of Sciences, Beijing 100190, China;5.5)Institute of Neuroscience, State Key Laboratory of Neuroscience, CAS Center for Excellence in Brain Science and Intelligence Technology, Chinese Academy of Sciences, Shanghai 200031, China;6.6)Shanghai Center for Brain Science and Brain-Inspired Intelligence Technology, Shanghai 201210, China;7.7)Shanghai Institute of Microsystem and Information Technology, Chinese Academy of Sciences, Shanghai 200050, China

the Strategic Priority Research Program of Chinese Academy of Science, Grant No.XDB32010100